Obtained B.Sc (Hons) and M.Sc. in Zoology from Bangalore University and Ph.D. in Molecular Biology from the
Indian Institute of Science,Bangalore,India. During the past 35 years he has established an active school
of research in the area of Animal Behaviour,Ecology and Evolution. The origin and evolution of cooperation
in animals,especially in social insects,such as ants,bees and wasps,is a major goal of his research. By
identifying and utilizing crucial elements in India’s biodiversity,he has added a special Indian flavour to
his research.